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Similarly we observed increased expression of miR-193a-5p in the tobacco treated cells, where as previous studies have shown miR-193a-5p is silenced by DNA hypermethylation in HNSCC and OSCC in smokers, indicating that the expression profile of miRNAs in oral cells may depend on the form of tobacco used 32 .We identified several miRNAs in chewing tobacco treated oral keratinocytes which are reported to be dysregulated in other cancer types but not in OSCC

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A separate meta-analysis found that paternal smoking was also associated with an increased risk of birth defects, specifically cardiac defects and orofacial defects

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